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Most species of ''Hygrocybe'' are soil-dwelling, though a few (such as ''Hygrocybe mexicana'' and ''H. rosea'') are only known from mossy tree trunks or logs.<ref name=Lodgeetal/> In Europe, species are typical of unimproved (nutrient-poor), short-sward grasslands, often termed "[[waxcap grassland]]s",<ref name="Rotheroe1996"/> but elsewhere they are more commonly found in woodland.
Their metabolism has long been debated, but recent research suggests that they are not [[saprotrophic]] <ref name="Seitzman" /><ref name=":0">{{Cite journal|last=Halbwachs|first=Hans|last2=Dentinger|first2=Bryn T.M.|last3=Detheridge|first3=Andrew P.|last4=Karasch|first4=Peter|last5=Griffith|first5=Gareth W.|date=December 2013|title=Hyphae of waxcap fungi colonise plant roots|url=https://doi.org/10.1016/j.funeco.2013.08.003|journal=Fungal Ecology|volume=6|issue=6|pages=487–492|doi=10.1016/j.funeco.2013.08.003|issn=1754-5048}}</ref> but rather [[symbiosis|symbiotically]] associated the roots of higher plants or mosses. Hyphae of ''[[Hygrocybe conica|H. conica]]'' have been detected in plant roots <ref name=":0" /> and other waxcap species have been detected as systemic endophytes of ''Plantago lanceolata''
Species are distributed worldwide, from the tropics to the sub-polar regions. Around 150 have been described to date.<ref name="DicFungi" />
==Conservation==
{{Main|Waxcap grassland}}
In Europe, waxcap grasslands and their associated fungi are of conservation concern, since unimproved grasslands (formerly commonplace) have declined dramatically as a result of changes in agricultural practice. As a result, by 1993, 89% of European ''Hygrocybe'' species appeared on one or more national [[red list]]s of threatened fungi.<ref name="Griffith2002"/>
In several countries, action has been taken to conserve waxcap grasslands and some of the rarer ''Hygrocybe'' species. Guidance
In the United Kingdom, some grasslands have gained a measure of legal protection as [[Sites of Special Scientific Interest]] because of their waxcap interest, notably [[Llanishen Reservoir]] in Cardiff, where the original [[SSSI]] notification by the [[Countryside Council for Wales]] in 2006 <ref>{{Cite news|url=http://archive.li/ZGZda|title=Llanishen reservoir action group news|date=2007-07-27|work=archive.li|access-date=2018-05-21}}</ref> was upheld following judicial review
==Economic usage==
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