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Grylloblattidae Temporal range: Triassic–Recent PreꞒ Ꞓ O S D C P T J K Pg N ^[1]

Galloisiana nipponensis
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Notoptera^[2]
Suborder:	Grylloblattodea
Family:	Grylloblattidae E. M. Walker, 1914
Genera
Galloisiana Grylloblatta Grylloblattella Grylloblattina Namkungia

Grylloblattidae is a family of extremophile (psychrophile) and wingless insects that live in the cold on top of mountains. It belongs, along with Mantophasmatidae, to the order Notoptera.^[2]

Overview

Galloisiana nipponensis

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Narwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

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Narwhals, Narwhals Swimming in the ocean Pretty big and pretty white They beat a polar bear in a fight

Like an underwater unicorn They've got a kick-ass facial horn They're the Jedi of the sea They stop Cthulu eating ye

Narwhals They are Narwhals Narwhals Just don't let 'em touch your balls Narwhals They are Narwhals Narwhals Inventors of the Shish Kebab

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(Narwhals, narwhals)

Narwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

Narwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

Narwhals Are really, really good You'll find them in the sea They're never in the woods Narwhals You know they're just so sweet They could stab you in the face With their long protruding teeth! Yeah!

It's a tooth A tooth, tooth baby They don't need a dentist Coz they're washin' it daily

It's a tooth! You know it's the truth! You don't need no more proof Coz I'm telling you it's a toothNarwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

Narwhals, Narwhals Swimming in the ocean Pretty big and pretty white They beat a polar bear in a fight

Like an underwater unicorn They've got a kick-ass facial horn They're the Jedi of the sea They stop Cthulu eating ye

Narwhals They are Narwhals Narwhals Just don't let 'em touch your balls Narwhals They are Narwhals Narwhals Inventors of the Shish Kebab

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(Narwhals, narwhals)

Narwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

Narwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

Narwhals Are really, really good You'll find them in the sea They're never in the woods Narwhals You know they're just so sweet They could stab you in the face With their long protruding teeth! Yeah!

It's a tooth A tooth, tooth baby They don't need a dentist Coz they're washin' it daily

It's a tooth! You know it's the truth! You don't need no more proof Coz I'm telling you it's a toothNarwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

Narwhals, Narwhals Swimming in the ocean Pretty big and pretty white They beat a polar bear in a fight

Like an underwater unicorn They've got a kick-ass facial horn They're the Jedi of the sea They stop Cthulu eating ye

Narwhals They are Narwhals Narwhals Just don't let 'em touch your balls Narwhals They are Narwhals Narwhals Inventors of the Shish Kebab

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(Narwhals, narwhals)

Narwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

Narwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

Narwhals Are really, really good You'll find them in the sea They're never in the woods Narwhals You know they're just so sweet They could stab you in the face With their long protruding teeth! Yeah!

It's a tooth A tooth, tooth baby They don't need a dentist Coz they're washin' it daily

It's a tooth! You know it's the truth! You don't need no more proof Coz I'm telling you it's a toothNarwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

Narwhals, Narwhals Swimming in the ocean Pretty big and pretty white They beat a polar bear in a fight

Like an underwater unicorn They've got a kick-ass facial horn They're the Jedi of the sea They stop Cthulu eating ye

Narwhals They are Narwhals Narwhals Just don't let 'em touch your balls Narwhals They are Narwhals Narwhals Inventors of the Shish Kebab

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(Narwhals, narwhals)

Narwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

Narwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

Narwhals Are really, really good You'll find them in the sea They're never in the woods Narwhals You know they're just so sweet They could stab you in the face With their long protruding teeth! Yeah!

It's a tooth A tooth, tooth baby They don't need a dentist Coz they're washin' it daily

It's a tooth! You know it's the truth! You don't need no more proof Coz I'm telling you it's a toothNarwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

Narwhals, Narwhals Swimming in the ocean Pretty big and pretty white They beat a polar bear in a fight

Like an underwater unicorn They've got a kick-ass facial horn They're the Jedi of the sea They stop Cthulu eating ye

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(Narwhals, narwhals)

Narwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

Narwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

Narwhals Are really, really good You'll find them in the sea They're never in the woods Narwhals You know they're just so sweet They could stab you in the face With their long protruding teeth! Yeah!

It's a tooth A tooth, tooth baby They don't need a dentist Coz they're washin' it daily

It's a tooth! You know it's the truth! You don't need no more proof Coz I'm telling you it's a toothNarwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

Narwhals, Narwhals Swimming in the ocean Pretty big and pretty white They beat a polar bear in a fight

Like an underwater unicorn They've got a kick-ass facial horn They're the Jedi of the sea They stop Cthulu eating ye

Narwhals They are Narwhals Narwhals Just don't let 'em touch your balls Narwhals They are Narwhals Narwhals Inventors of the Shish Kebab

Narwhals They are Narwhals Narwhals Just don't let 'em touch your balls Narwhals They are Narwhals Narwhals

(Narwhals, narwhals)

Narwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

Narwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

Narwhals Are really, really good You'll find them in the sea They're never in the woods Narwhals You know they're just so sweet They could stab you in the face With their long protruding teeth! Yeah!

It's a tooth A tooth, tooth baby They don't need a dentist Coz they're washin' it daily

It's a tooth! You know it's the truth! You don't need no more proof Coz I'm telling you it's a toothNarwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

Narwhals, Narwhals Swimming in the ocean Pretty big and pretty white They beat a polar bear in a fight

Like an underwater unicorn They've got a kick-ass facial horn They're the Jedi of the sea They stop Cthulu eating ye

Narwhals They are Narwhals Narwhals Just don't let 'em touch your balls Narwhals They are Narwhals Narwhals Inventors of the Shish Kebab

Narwhals They are Narwhals Narwhals Just don't let 'em touch your balls Narwhals They are Narwhals Narwhals

(Narwhals, narwhals)

Narwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

Narwhals, Narwhals Swimming in the ocean Causing a commotion Coz they are so awesome

Narwhals Are really, really good You'll find them in the sea They're never in the woods Narwhals You know they're just so sweet They could stab you in the face With their long protruding teeth! Yeah!

It's a tooth A tooth, tooth baby They don't need a dentist Coz they're washin' it daily

It's a tooth! You know it's the truth! You don't need no more proof Coz I'm telling you it's a tooth |doi=10.1016/j.ympev.2005.09.020 |pmid=16321547 |issue=1}}</ref>

The family is placed in its own suborder, Grylloblattodea, and in its own former^[2] order, Grylloblattaria. It contains 5 genera and 34 species.^[3]^[4]

Habitat

Grylloblattodea are nocturnal extremophiles typically found in leaf litter and under stones in extremely cold environments, usually at higher elevations. They are known to inhabit cold temperate forests to glaciers and ice sheets. Their optimal living temperature is between 1-4 °C (33.8-39.2 °F). They can be killed at colder temperatures due to ice formation in the body, so when the temperature drops below their optimal range they survive by living under snow pack near the soil.^[5]

Diet

They are omnivorous, but feed primarily on deceased arthropods. When arthropod carcasses are scarce, their diet relies heavily on plant material.^[5]

References

^ H. V. Hoell, J. T. Doyen & A. H. Purcell (1998). Introduction to Insect Biology and Diversity (2nd ed.). Oxford University Press. p. 320. ISBN 0-19-510033-6.
^ ^a ^b ^c Arillo, A. & M. Engel (2006) Rock Crawlers in Baltic Amber (Notoptera: Mantophasmatodea). American Museum Novitates 3539:1-10
^ Terry L. Erwin (1997). "Biodiversity at its utmost: tropical forest beetles". In Marjorie L. Reaka-Kudla, Don E. Wilson & Edward O. Wilson (ed.). Biodiversity II. Washington, D.C.: Joseph Henry Press. pp. 27–40. ISBN 978-0-309-05584-0.
^ Zhang, Z.-Q. (2011). "Phylum Arthropoda von Siebold, 1848 In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness" (PDF). Zootaxa. 3148: 99–103.
^ ^a ^b David Grimaldi, Michael S. Engel (2005). "Polyneoptera: Grylloblattodea: The Ice Crawlers". Evolution of the Insects. New York City: Cambridge University Press. pp. 222–224. ISBN 9780521821490.

External links

Media related to Grylloblattidae at Wikimedia Commons
Data related to Grylloblattidae at Wikispecies
Video of a walking grylloblattid

v t e Extant Notoptera families
Kingdom: Animalia Phylum: Arthropoda Class: Insecta Subclass: Pterygota Infraclass: Neoptera
Grylloblattodea	Grylloblattidae (ice crawlers or icebugs)
Mantophasmatodea	Mantophasmatidae (rock crawlers or gladiators)

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